Do Snake Venoms Evolve According to the Prey They Are Intended to Kill, Or Do Snakes Just Dial Up the Dosage or Concentration of Whatever Venom They Have to the Size of the Prey in their Sights?

Snake venoms are overwhelmingly composed  of dissolved protein  mixtures which typically exert their action on a victim in one of the following three ways, and sometimes in combination:

·        Hemotoxins. Depending on their mix of proteins, these can either cause the blood to clot precipitously, causing cardiovascular blockages (essentially heart attacks) in extreme cases, or alternatively, cause it to thin so quickly that the victim bleeds out.  This is one of the  modes of action of rattlesnake venoms.

 

·        Neurotoxins.  These disrupt either or both the voluntary and autonomic nervous system of the victim, rendering it incapable of escape, and in extreme cases, cause the victim to stop breathing or to convulse. This is the primary  mode of action of cobra venoms.

 

·         Tissue Necrotizers:  These basically break down the muscular and connective tissue at the site of the bite, and basically cause an ever expanding dead zone until the toxin is so diluted as to lose effect. Often the victim dies before this can happen. This is the mode of action of the fer-de-lance the most prominent perpetrator of poisonous snake bites in Latin America.

The popularity of proteomics and the wider availability of its instrumentation  leads to “venomics” & confirms evolutionary relationships & distinctions  among the many differing families  of poisonous snakes

 

Can you tell a venomous snake just by a chemical analysis of its venom?

The short, and historically most orthodox answer,  is, yes.

For most of the history of snake venom studies , the relatively small number of total analyses achieved indicated that most snakes tend to have signature venom protein mixes.

Now, given the widespread use of automated proteomics analyzers and advances in protein sequence informatics, much of this  old work is being confirmed with more  complete analyses of more venoms from more snakes and especially from differing families of snakes. Indeed, you can now search a variety of biomedical, chemical and pharmaceutical  databases under “Venomics” for on-point papers.

 And the new results can be extended to say that snakes that are in the same genus tend to have rather similar, if not always quite identical, cocktails of toxins.

Such variations as are found, are thought to be minor,  the results of random genetic drift, and to be incidental to the  study of  either the lineage, the  or day-to-day functioning of the snake.

There is  some thought that evolutionary relationships suggested by traditional morphological characters or modern genomics ,  can predict,  with reasonable certainty what venom toxins given families of snakes are likely to share, even before all their members are tested.

As a kind of real-world proof of the all-in-the-family nature  of protein toxins , consider that snake antivenoms (which work on biochemical recognition and disabling of venom proteins through binding them up)  work best when customized to the type of specific snake that does the biting. Such antitoxins are called monovalents, and are the treatment of choice whenever available.

But  given that maintaining a large and varied collection of monovalent antivenoms  is not always practical in areas where there are many possible different poisonous snakes which bite, some polyvalent vaccines have had to be developed which counteract, to some degree,  venoms from a variety of unrelated species.

Despite their polyvalent intent, their comparative effectiveness is positively correlated  with snakebites  that are evolutionarily related to the species from which the original pharmaceutical venom/vaccine was drawn.  In other words, antivenom that derives from a specific type of cobra,  is likely to work better with other persons bitten by somewhat different cobras, than it is to work on people bitten by rattlesnakes.

The prevalent theory is that the biosynthesis of toxins is likely to be highly conserved  over many generations and over differing populations of snake families  because their venom seems so central to the function of the snake, and that even incidental variations in toxins caused my random mutations could come at a cost to the snake in reduced prey killing capacity.

But  the dismissal of the significance of small differences in toxin blends within related snakes  is now being challenged,  and variations are being seen as having some meaningful function, and not just being blips in encoding that have no real life meaning to the snake.

Why have differing toxins within the same family of snakes? Couldn’t the snake just adjust the dosage to the size of the prey?

Differing types of snakes have differing quantities of venom available for hunting or defense, but certain general rules can predict among populations of the same snakes which will have more venom on average.

Generally, the older the snake of a given species is , the larger it is, and therefore the larger its storage and potential supply of venom for “envenomation, ”  when compared to younger and smaller snakes of the same species.

Furthermore female snakes generally have significantly more venom than males for two  readily  understood reasons: They are generally larger than males and have larger heads, and the head is where most of the venom production and storage is confined in all but a tiny handful of snake species.

But, it turns out that many adult snakes, whether large or small,  are actually thrifty in their use of venom.

They adjust the amount of venom to the size of their prey.  A mouse might get less venom per bite  than a gopher, for example.

Although as a general rule, given a choice between small and large prey , larger snakes will choose larger prey, and are willing to “spend the venom”  in order to get the larger meal.

The saw-scaled viper family and their venoms

A particularly  interesting comparison test of the venoms of four kinds of  saw-scaled vipers done by Barlow et al. (2009, cited below).

These four related viper species  are a genuine menace to public health in that they cause the majority of snakebite cases in North Africa and the adjacent Middle East.

 (The fact that some of them kill harmful vermin, may, of course be an offsetting factor, before deciding that they all should be wiped out indiscriminately.)

A comparison of the  electrophoretic separations of major toxin proteins  from the four different types of saw-scaled viper  showed that they were indeed, matching in the major protein bands,  something predictable owing to conventional thinking about the molecular evolution of snake venoms.

But there was a surprising amount of variation in the minor bands.

And this was also reflected in the fact that in the Middle East and in Northern Africa, saw scale viper antivenoms were markedly more effective against  Middle Eastern saw scale viper bites than they were against the bites of  the North African saw scale vipers that were presumed to be their evolutionarily closest relatives.

The drop off in effectiveness is notable in clinical terms and makes follow-up injections or IV drips of antivenom necessary more often in North Africa than they were in the Middle East.

A question arose:  Beyond any clinical significance, did these  variant protein toxins  have a purpose in the life of the snake?  Or, were they just the result of geographic isolation and random genetic drift?

Prey size vs. prey type: Which  plays a bigger role in the  evolution of venoms?

One competing theory of comparative venom toxicity  is that snakes within the same general family of snakes that prey on larger animals will eventually evolve   not only to produce more of the same general type of venom,  but also evolve higher, and therefore more potent concentrations of venom, with which to immobilize and kill their “big game.”

Its principal rival theory  is that differing prey  specialists within the same general family of snakes are more likely to require,  and eventually develop over succeeding generations venoms that gradually become more distinct, depending for success in killing prey, not so much on increasing the amount or the concentration of venom injected,   but rather on matching  the prey’s particular susceptibility to the venom.

Saw scale vipers represent a wonderful set of relatives upon which to test these competing outlooks.

Several types of saw scaled vipers have become small mammal specialists (usually eating rodents)  in terms of their prey.  Most of these are in the Middle East.

Others have a mixed diet of rodents and invertebrates.

(Most commonly in the Middle East and Africa, the invertebrates are various types of scorpions. )

Still others are almost exclusively scorpion feeders. These vipers are found only in North Africa.

It should be understood that even the biggest scorpions are but a fraction of the size and weight of the mice and rats typically consumed by either the rodent specialist,  or the mixed-diet-consuming saw scale  vipers.

Consequently “big game venom” should actually be more toxic to little game, because of its overall increased potency.

In other words, it may not be necessary to shoot a big-game venom cannon ball to kill a scorpion, but if it were to happen,  the cannon ball should surely be lethal to the scorpion.

Using ingenious methods, such as directly injecting carefully calibrated, matching fluid amounts of venom directly into  test scorpions from all four types of saw scale viper, and then seeing which venoms were, “ounce for ounce” more deadly, the team found something unexpected.

The “big game hunters”  of rats and mice, who would presumably need more toxicity to bring down their big game, actually  had venom that was notably less toxic to the small size game of scorpions, than the venom   of the mixed-bag-diet hunters.

And even more  intriguingly,   the mixed-bag-hunter venom was still less toxic to the scorpions than that of the specialized scorpion hunters.

The most toxic & effective venoms against the small game of scorpions, came from scorpion specialists, not from big-game or mixed-game specialists, who routinely had to knock down and kill bigger game.

In other words, the venom of North African scorpion specialists had become distinct enough through evolutionary adaptation to its prey, to make a difference in the life of its particular type of saw scale viper.

And maybe that distinction was now big enough to explain why Middle Eastern antivenom vaccines against North African saw scale vipers were seeing decreased efficiency.

Are the prey utterly without their own evolutionary defenses against specially  evolved snake venom?

This would seem to be the case, but in a number of settings, various ground squirrels, gophers and badgers, whose paths frequently intersect with predatory snakes like rattlers, seem to have grown immune to the local venom (see, for example Hayes, Lavin-Murcio & Kardong, 1995, cited below). This is also the case with some eels that live alongside sea snakes that hunt for eels (Heatwole & Pran, 1995).

Whether or not this leads to the evolutionary  alteration of the local snake venom to make if effective once again, has not yet been studied.

But in a particularly intriguing case, a poisonous sea snake that has become a fish egg eating specialists, and therefore has no need to immobilize and kill its prey (the eggs don’t run away and are not guarded by their parent fish) has actually evolved venom of greatly reduced toxicity, seeming to spare itself unnecessary toxin protein production (Li, Fry & Manjunatha Kini, 2005).

 

Tony Stankus [email protected] Life Sciences Librarian & Professor

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Thanks.| {It is|It's} {appropriate|perfect|the best} time to make some plans for the future and {it is|it's} time to be happy. {I have|I've} read this post and if I could I {want to|wish to|desire to} suggest you {few|some} interesting things or {advice|suggestions|tips}. {Perhaps|Maybe} you {could|can} write next articles referring to this article. I {want to|wish to|desire to} read {more|even more} things about it!| {It is|It's} {appropriate|perfect|the best} time to make {a few|some} plans for {the future|the longer term|the long run} and {it is|it's} time to be happy. {I have|I've} {read|learn} this {post|submit|publish|put up} and if I {may just|may|could} I {want to|wish to|desire to} {suggest|recommend|counsel} you {few|some} {interesting|fascinating|attention-grabbing} {things|issues} or {advice|suggestions|tips}. {Perhaps|Maybe} you {could|can} write {next|subsequent} articles {relating to|referring to|regarding} this article. I {want to|wish to|desire to} {read|learn} {more|even more} {things|issues} {approximately|about} it!| {I have|I've} been {surfing|browsing} {online|on-line} {more than|greater than} {three|3} hours {these days|nowadays|today|lately|as of late}, {yet|but} I {never|by no means} {found|discovered} any {interesting|fascinating|attention-grabbing} article like yours. {It's|It is} {lovely|pretty|beautiful} {worth|value|price} {enough|sufficient} for me. {In my opinion|Personally|In my view}, if all {webmasters|site owners|website owners|web owners} and bloggers made {just right|good|excellent} {content|content material} as {you did|you probably did}, the {internet|net|web} {will be|shall be|might be|will probably be|can be|will likely be} {much more|a lot more} {useful|helpful} than ever before.| Ahaa, its {nice|pleasant|good|fastidious} {discussion|conversation|dialogue} {regarding|concerning|about|on the topic of} this {article|post|piece of writing|paragraph} {here|at this place} at this {blog|weblog|webpage|website|web site}, I have read all that, so {now|at this time} me also commenting {here|at this place}.| I am sure this {article|post|piece of writing|paragraph} has touched all the internet {users|people|viewers|visitors}, its really really {nice|pleasant|good|fastidious} {article|post|piece of writing|paragraph} on building up new {blog|weblog|webpage|website|web site}.| Wow, this {article|post|piece of writing|paragraph} is {nice|pleasant|good|fastidious}, my {sister|younger sister} is analyzing {such|these|these kinds of} things, {so|thus|therefore} I am going to {tell|inform|let know|convey} her.| {Saved as a favorite|bookmarked!!}, {I really like|I like|I love} {your blog|your site|your web site|your website}!| Way cool! Some {very|extremely} valid points! I appreciate you {writing this|penning this} {article|post|write-up} {and the|and also the|plus the} rest of the {site is|website is} {also very|extremely|very|also really|really} good.| Hi, {I do believe|I do think} {this is an excellent|this is a great} {blog|website|web site|site}. I stumbledupon it ;) {I will|I am going to|I'm going to|I may} {come back|return|revisit} {once again|yet again} {since I|since i have} {bookmarked|book marked|book-marked|saved as a favorite} it. Money and freedom {is the best|is the greatest} way to change, may you be rich and continue to {help|guide} {other people|others}.| Woah! I'm really {loving|enjoying|digging} the template/theme of this {site|website|blog}. It's simple, yet effective. A lot of times it's {very hard|very difficult|challenging|tough|difficult|hard} to get that "perfect balance" between {superb usability|user friendliness|usability} and {visual appearance|visual appeal|appearance}. I must say {that you've|you have|you've} done a {awesome|amazing|very good|superb|fantastic|excellent|great} job with this. {In addition|Additionally|Also}, the blog loads {very|extremely|super} {fast|quick} for me on {Safari|Internet explorer|Chrome|Opera|Firefox}. {Superb|Exceptional|Outstanding|Excellent} Blog!| These are {really|actually|in fact|truly|genuinely} {great|enormous|impressive|wonderful|fantastic} ideas in {regarding|concerning|about|on the topic of} blogging. You have touched some {nice|pleasant|good|fastidious} {points|factors|things} here. Any way keep up wrinting.| {I love|I really like|I enjoy|I like|Everyone loves} what you guys {are|are usually|tend to be} up too. {This sort of|This type of|Such|This kind of} clever work and {exposure|coverage|reporting}! Keep up the {superb|terrific|very good|great|good|awesome|fantastic|excellent|amazing|wonderful} works guys I've {incorporated||added|included} you guys to {|my|our||my personal|my own} blogroll.| {Howdy|Hi there|Hey there|Hi|Hello|Hey}! Someone in my {Myspace|Facebook} group shared this {site|website} with us so I came to {give it a look|look it over|take a look|check it out}. I'm definitely {enjoying|loving} the information. I'm {book-marking|bookmarking} and will be tweeting this to my followers! {Terrific|Wonderful|Great|Fantastic|Outstanding|Exceptional|Superb|Excellent} blog and {wonderful|terrific|brilliant|amazing|great|excellent|fantastic|outstanding|superb} {style and design|design and style|design}.| {I love|I really like|I enjoy|I like|Everyone loves} what you guys {are|are usually|tend to be} up too. {This sort of|This type of|Such|This kind of} clever work and {exposure|coverage|reporting}! Keep up the {superb|terrific|very good|great|good|awesome|fantastic|excellent|amazing|wonderful} works guys I've {incorporated|added|included} you guys to {|my|our|my personal|my own} blogroll.| {Howdy|Hi there|Hey there|Hi|Hello|Hey} would you mind {stating|sharing} which blog platform you're {working with|using}? I'm {looking|planning|going} to start my own blog {in the near future|soon} but I'm having a {tough|difficult|hard} time {making a decision|selecting|choosing|deciding} between BlogEngine/Wordpress/B2evolution and Drupal. The reason I ask is because your {design and style|design|layout} seems different then most blogs and I'm looking for something {completely unique|unique}. P.S {My apologies|Apologies|Sorry} for {getting|being} off-topic but I had to ask!| {Howdy|Hi there|Hi|Hey there|Hello|Hey} would you mind letting me know which {webhost|hosting company|web host} you're {utilizing|working with|using}? I've loaded your blog in 3 {completely different|different} {internet browsers|web browsers|browsers} and I must say this blog loads a lot {quicker|faster} then most. Can you {suggest|recommend} a good {internet hosting|web hosting|hosting} provider at a {honest|reasonable|fair} price? {Thanks a lot|Kudos|Cheers|Thank you|Many thanks|Thanks}, I appreciate it!| {I love|I really like|I like|Everyone loves} it {when people|when individuals|when folks|whenever people} {come together|get together} and share {opinions|thoughts|views|ideas}. Great {blog|website|site}, {keep it up|continue the good work|stick with it}!| Thank you for the {auspicious|good} writeup. It in fact was a amusement account it. Look advanced to {far|more} added agreeable from you! {By the way|However}, how {can|could} we communicate?| {Howdy|Hi there|Hey there|Hello|Hey} just wanted to give you a quick heads up. The {text|words} in your {content|post|article} seem to be running off the screen in {Ie|Internet explorer|Chrome|Firefox|Safari|Opera}. I'm not sure if this is a {format|formatting} issue or something to do with {web browser|internet browser|browser} compatibility but I {thought|figured} I'd post to let you know. The {style and design|design and style|layout|design} look great though! Hope you get the {problem|issue} {solved|resolved|fixed} soon. {Kudos|Cheers|Many thanks|Thanks}| This is a topic {that is|that's|which is} {close to|near to} my heart... {Cheers|Many thanks|Best wishes|Take care|Thank you}! {Where|Exactly where} are your contact details though?| It's very {easy|simple|trouble-free|straightforward|effortless} to find out any {topic|matter} on {net|web} as compared to {books|textbooks}, as I found this {article|post|piece of writing|paragraph} at this {website|web site|site|web page}.| Does your {site|website|blog} have a contact page? I'm having {a tough time|problems|trouble} locating it but, I'd like to {send|shoot} you an {e-mail|email}. I've got some {creative ideas|recommendations|suggestions|ideas} for your blog you might be interested in hearing. Either way, great {site|website|blog} and I look forward to seeing it {develop|improve|expand|grow} over time.| {Hola|Hey there|Hi|Hello|Greetings}! I've been {following|reading} your {site|web site|website|weblog|blog} for {a long time|a while|some time} now and finally got the {bravery|courage} to go ahead and give you a shout out from {New Caney|Kingwood|Huffman|Porter|Houston|Dallas|Austin|Lubbock|Humble|Atascocita} {Tx|Texas}! Just wanted to {tell you|mention|say} keep up the {fantastic|excellent|great|good} {job|work}!| Greetings from {Idaho|Carolina|Ohio|Colorado|Florida|Los angeles|California}! I'm {bored to tears|bored to death|bored} at work so I decided to {check out|browse} your {site|website|blog} on my iphone during lunch break. I {enjoy|really like|love} the {knowledge|info|information} you {present|provide} here and can't wait to take a look when I get home. I'm {shocked|amazed|surprised} at how {quick|fast} your blog loaded on my {mobile|cell phone|phone} .. I'm not even using WIFI, just 3G .. {Anyhow|Anyways}, {awesome|amazing|very good|superb|good|wonderful|fantastic|excellent|great} {site|blog}!| Its {like you|such as you} {read|learn} my {mind|thoughts}! You {seem|appear} {to understand|to know|to grasp} {so much|a lot} {approximately|about} this, {like you|such as you} wrote the {book|e-book|guide|ebook|e book} in it or something. {I think|I feel|I believe} {that you|that you simply|that you just} {could|can} do with {some|a few} {%|p.c.|percent} to {force|pressure|drive|power} the message {house|home} {a bit|a little bit}, {however|but} {other than|instead of} that, {this is|that is} {great|wonderful|fantastic|magnificent|excellent} blog. {A great|An excellent|A fantastic} read. {I'll|I will} {definitely|certainly} be back.| I visited {multiple|many|several|various} {websites|sites|web sites|web pages|blogs} {but|except|however} the audio {quality|feature} for audio songs {current|present|existing} at this {website|web site|site|web page} is {really|actually|in fact|truly|genuinely} {marvelous|wonderful|excellent|fabulous|superb}.| {Howdy|Hi there|Hi|Hello}, i read your blog {occasionally|from time to time} and i own a similar one and i was just {wondering|curious} if you get a lot of spam {comments|responses|feedback|remarks}? If so how do you {prevent|reduce|stop|protect against} it, any plugin or anything you can {advise|suggest|recommend}? I get so much lately it's driving me {mad|insane|crazy} so any {assistance|help|support} is very much appreciated.| Greetings! {Very helpful|Very useful} advice {within this|in this particular} {article|post}! {It is the|It's the} little changes {that make|which will make|that produce|that will make} {the biggest|the largest|the greatest|the most important|the most significant} changes. {Thanks a lot|Thanks|Many thanks} for sharing!| {I really|I truly|I seriously|I absolutely} love {your blog|your site|your website}.. {Very nice|Excellent|Pleasant|Great} colors & theme. Did you {create|develop|make|build} {this website|this site|this web site|this amazing site} yourself? Please reply back as I'm {looking to|trying to|planning to|wanting to|hoping to|attempting to} create {my own|my very own|my own personal} {blog|website|site} and {would like to|want to|would love to} {know|learn|find out} where you got this from or {what the|exactly what the|just what the} theme {is called|is named}. {Thanks|Many thanks|Thank you|Cheers|Appreciate it|Kudos}!| {Hi there|Hello there|Howdy}! This {post|article|blog post} {couldn't|could not} be written {any better|much better}! {Reading through|Looking at|Going through|Looking through} this {post|article} reminds me of my previous roommate! He {always|constantly|continually} kept {talking about|preaching about} this. {I will|I'll|I am going to|I most certainly will} {forward|send} {this article|this information|this post} to him. {Pretty sure|Fairly certain} {he will|he'll|he's going to} {have a good|have a very good|have a great} read. {Thank you for|Thanks for|Many thanks for|I appreciate you for} sharing!| {Wow|Whoa|Incredible|Amazing}! This blog looks {exactly|just} like my old one! It's on a {completely|entirely|totally} different {topic|subject} but it has pretty much the same {layout|page layout} and design. {Excellent|Wonderful|Great|Outstanding|Superb} choice of colors!| {There is|There's} {definately|certainly} {a lot to|a great deal to} {know about|learn about|find out about} this {subject|topic|issue}. {I like|I love|I really like} {all the|all of the} points {you made|you've made|you have made}.| {You made|You've made|You have made} some {decent|good|really good} points there. I {looked|checked} {on the internet|on the web|on the net} {for more info|for more information|to find out more|to learn more|for additional information} about the issue and found {most individuals|most people} will go along with your views on {this website|this site|this web site}.| {Hi|Hello|Hi there|What's up}, I {log on to|check|read} your {new stuff|blogs|blog} {regularly|like every week|daily|on a regular basis}. Your {story-telling|writing|humoristic} style is {awesome|witty}, keep {doing what you're doing|up the good work|it up}!| I {simply|just} {could not|couldn't} {leave|depart|go away} your {site|web site|website} {prior to|before} suggesting that I {really|extremely|actually} {enjoyed|loved} {the standard|the usual} {information|info} {a person|an individual} {supply|provide} {for your|on your|in your|to your} {visitors|guests}? Is {going to|gonna} be {back|again} {frequently|regularly|incessantly|steadily|ceaselessly|often|continuously} {in order to|to} {check up on|check out|inspect|investigate cross-check} new posts| {I wanted|I needed|I want to|I need to} to thank you for this {great|excellent|fantastic|wonderful|good|very good} read!! I {definitely|certainly|absolutely} {enjoyed|loved} every {little bit of|bit of} it. {I have|I've got|I have got} you {bookmarked|book marked|book-marked|saved as a favorite} {to check out|to look at} new {stuff you|things you} post…| {Hi|Hello|Hi there|What's up}, just wanted to {mention|say|tell you}, I {enjoyed|liked|loved} this {article|post|blog post}. It was {inspiring|funny|practical|helpful}. Keep on posting!| I {{leave|drop|{write|create}} a {comment|leave a response}|drop a {comment|leave a response}|{comment|leave a response}} {each time|when|whenever} I {appreciate|like|especially enjoy} a {post|article} on a {site|{blog|website}|site|website} or {I have|if I have} something to {add|contribute|valuable to contribute} {to the discussion|to the conversation}. {It is|Usually it is|Usually it's|It's} {a result of|triggered by|caused by} the {passion|fire|sincerness} {communicated|displayed} in the {post|article} I {read|looked at|browsed}. And {on|after} this {post|article} SLA Biomedical & Life Sciences Division Blog: Do Snake Venoms Evolve According to the Prey They Are Intended to Kill, Or Do Snakes Just Dial Up the Dosage or Concentration of Whatever Venom They Have to the Size of the Prey in their Sights?. I {{was|was actually} moved|{was|was actually} excited} enough to {drop|{leave|drop|{write|create}}|post} a {thought|{comment|{comment|leave a response}a response}} {:-P|:)|;)|;-)|:-)} I {do have|actually do have} {{some|a few} questions|a couple of questions|2 questions} for you {if you {don't|do not|usually do not|tend not to} mind|if it's {allright|okay}}. {Is it|Could it be} {just|only|simply} me or {do|does it {seem|appear|give the impression|look|look as if|look like} like} {some|a few} of {the|these} {comments|responses|remarks} {look|appear|come across} {like they are|as if they are|like} {coming from|written by|left by} brain dead {people|visitors|folks|individuals}? :-P And, if you are {posting|writing} {on|at} {other|additional} {sites|social sites|online sites|online social sites|places}, {I'd|I would} like to {follow|keep up with} {you|{anything|everything} {new|fresh} you have to post}. {Could|Would} you {list|make a list} {all|every one|the complete urls} of {your|all your} {social|communal|community|public|shared} {pages|sites} like your {twitter feed, Facebook page or linkedin profile|linkedin profile, Facebook page or twitter feed|Facebook page, twitter feed, or linkedin profile}?| {Hi there|Hello}, I enjoy reading {all of|through} your {article|post|article post}. I {like|wanted} to write a little comment to support you.| I {always|constantly|every time} spent my half an hour to read this {blog|weblog|webpage|website|web site}'s {articles|posts|articles or reviews|content} {everyday|daily|every day|all the time} along with a {cup|mug} of coffee.| I {always|for all time|all the time|constantly|every time} emailed this {blog|weblog|webpage|website|web site} post page to all my {friends|associates|contacts}, {because|since|as|for the reason that} if like to read it {then|after that|next|afterward} my {friends|links|contacts} will too.| My {coder|programmer|developer} is trying to {persuade|convince} me to move to .net from PHP. I have always disliked the idea because of the {expenses|costs}. But he's tryiong none the less. I've been using {Movable-type|WordPress} on {a number of|a variety of|numerous|several|various} websites for about a year and am {nervous|anxious|worried|concerned} about switching to another platform. I have heard {fantastic|very good|excellent|great|good} things about blogengine.net. Is there a way I can {transfer|import} all my wordpress {content|posts} into it? {Any kind of|Any} help would be {really|greatly} appreciated!| {Hello|Hi|Hello there|Hi there|Howdy|Good day}! I could have sworn I've {been to|visited} {this blog|this web site|this website|this site|your blog} before but after {browsing through|going through|looking at} {some of the|a few of the|many of the} {posts|articles} I realized it's new to me. {Anyways|Anyhow|Nonetheless|Regardless}, I'm {definitely|certainly} {happy|pleased|delighted} {I found|I discovered|I came across|I stumbled upon} it and I'll be {bookmarking|book-marking} it and checking back {frequently|regularly|often}!| {Terrific|Great|Wonderful} {article|work}! {This is|That is} {the type of|the kind of} {information|info} {that are meant to|that are supposed to|that should} be shared {around the|across the} {web|internet|net}. {Disgrace|Shame} on {the {seek|search} engines|Google} for {now not|not|no longer} positioning this {post|submit|publish|put up} {upper|higher}! Come on over and {talk over with|discuss with|seek advice from|visit|consult with} my {site|web site|website} . {Thank you|Thanks} =)| Heya {i'm|i am} for the first time here. I {came across|found} this board and I find It {truly|really} useful & it helped me out {a lot|much}. I hope to give something back and {help|aid} others like you {helped|aided} me.| {Hi|Hello|Hi there|Hello there|Howdy|Greetings}, {I think|I believe|I do believe|I do think|There's no doubt that} {your site|your website|your web site|your blog} {might be|may be|could be|could possibly be} having {browser|internet browser|web browser} compatibility {issues|problems}. {When I|Whenever I} {look at your|take a look at your} {website|web site|site|blog} in Safari, it looks fine {but when|however when|however, if|however, when} opening in {Internet Explorer|IE|I.E.}, {it has|it's got} some overlapping issues. {I just|I simply|I merely} wanted to {give you a|provide you with a} quick heads up! {Other than that|Apart from that|Besides that|Aside from that}, {fantastic|wonderful|great|excellent} {blog|website|site}!| {A person|Someone|Somebody} {necessarily|essentially} {lend a hand|help|assist} to make {seriously|critically|significantly|severely} {articles|posts} {I would|I might|I'd} state. {This is|That is} the {first|very first} time I frequented your {web page|website page} and {to this point|so far|thus far|up to now}? I {amazed|surprised} with the {research|analysis} you made to {create|make} {this actual|this particular} {post|submit|publish|put up} {incredible|amazing|extraordinary}. {Great|Wonderful|Fantastic|Magnificent|Excellent} {task|process|activity|job}!| Heya {i'm|i am} for {the primary|the first} time here. I {came across|found} this board and I {in finding|find|to find} It {truly|really} {useful|helpful} & it helped me out {a lot|much}. {I am hoping|I hope|I'm hoping} {to give|to offer|to provide|to present} {something|one thing} {back|again} and {help|aid} others {like you|such as you} {helped|aided} me.| {Hello|Hi|Hello there|Hi there|Howdy|Good day|Hey there}! {I just|I simply} {would like to|want to|wish to} {give you a|offer you a} {huge|big} thumbs up {for the|for your} {great|excellent} {info|information} {you have|you've got|you have got} {here|right here} on this post. {I will be|I'll be|I am} {coming back to|returning to} {your blog|your site|your website|your web site} for more soon.| I {always|all the time|every time} used to {read|study} {article|post|piece of writing|paragraph} in news papers but now as I am a user of {internet|web|net} {so|thus|therefore} from now I am using net for {articles|posts|articles or reviews|content}, thanks to web.| Your {way|method|means|mode} of {describing|explaining|telling} {everything|all|the whole thing} in this {article|post|piece of writing|paragraph} is {really|actually|in fact|truly|genuinely} {nice|pleasant|good|fastidious}, {all|every one} {can|be able to|be capable of} {easily|without difficulty|effortlessly|simply} {understand|know|be aware of} it, Thanks a lot.| {Hi|Hello} there, {I found|I discovered} your {blog|website|web site|site} {by means of|via|by the use of|by way of} Google {at the same time as|whilst|even as|while} {searching for|looking for} a {similar|comparable|related} {topic|matter|subject}, your {site|web site|website} {got here|came} up, it {looks|appears|seems|seems to be|appears to be like} {good|great}. {I have|I've} bookmarked it in my google bookmarks. {Hello|Hi} there, {simply|just} {turned into|became|was|become|changed into} {aware of|alert to} your {blog|weblog} {thru|through|via} Google, {and found|and located} that {it is|it's} {really|truly} informative. {I'm|I am} {gonna|going to} {watch out|be careful} for brussels. {I will|I'll} {appreciate|be grateful} {if you|should you|when you|in the event you|in case you|for those who|if you happen to} {continue|proceed} this {in future}. {A lot of|Lots of|Many|Numerous} {other folks|folks|other people|people} {will be|shall be|might be|will probably be|can be|will likely be} benefited {from your|out of your} writing. Cheers!| {I am|I'm} curious to find out what blog {system|platform} {you have been|you happen to be|you are|you're} {working with|utilizing|using}? I'm {experiencing|having} some {minor|small} security {problems|issues} with my latest {site|website|blog} and {I would|I'd} like to find something more {safe|risk-free|safeguarded|secure}. Do you have any {solutions|suggestions|recommendations}?| {I am|I'm} {extremely|really} impressed with your writing skills {and also|as well as} with the layout on your {blog|weblog}. Is this a paid theme or did you {customize|modify} it yourself? {Either way|Anyway} keep up the {nice|excellent} quality writing, {it's|it is} rare to see a {nice|great} blog like this one {these days|nowadays|today}.| {I am|I'm} {extremely|really} {inspired|impressed} {with your|together with your|along with your} writing {talents|skills|abilities} {and also|as {smartly|well|neatly} as} with the {layout|format|structure} {for your|on your|in your|to your} {blog|weblog}. {Is this|Is that this} a paid {subject|topic|subject matter|theme} or did you {customize|modify} it {yourself|your self}? {Either way|Anyway} {stay|keep} up the {nice|excellent} {quality|high quality} writing, {it's|it is} {rare|uncommon} {to peer|to see|to look} a {nice|great} {blog|weblog} like this one {these days|nowadays|today}..| {Hi|Hello}, Neat post. {There is|There's} {a problem|an issue} {with your|together with your|along with your} {site|web site|website} in {internet|web} explorer, {may|might|could|would} {check|test} this? IE {still|nonetheless} is the {marketplace|market} {leader|chief} and {a large|a good|a big|a huge} {part of|section of|component to|portion of|component of|element of} {other folks|folks|other people|people} will {leave out|omit|miss|pass over} your {great|wonderful|fantastic|magnificent|excellent} writing {due to|because of} this problem.| {I'm|I am} not sure where {you are|you're} getting your {info|information}, but {good|great} topic. I needs to spend some time learning {more|much more} or understanding more. Thanks for {great|wonderful|fantastic|magnificent|excellent} {information|info} I was looking for this {information|info} for my mission.| {Hi|Hello}, i think that i saw you visited my {blog|weblog|website|web site|site} {so|thus} i came to “return the favor”.{I am|I'm} {trying to|attempting to} find things to {improve|enhance} my {website|site|web site}!I suppose its ok to use {some of|a few of} your ideas!!\

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{ {I have|I've} been {surfing|browsing} online more than {three|3|2|4} hours today, yet I never found any interesting article like yours. {It's|It is} pretty worth enough for me. {In my opinion|Personally|In my view}, if all {webmasters|site owners|website owners|web owners} and bloggers made good content as you did, the {internet|net|web} will be {much more|a lot more} useful than ever before.| I {couldn't|could not} {resist|refrain from} commenting. {Very well|Perfectly|Well|Exceptionally well} written!| {I will|I'll} {right away|immediately} {take hold of|grab|clutch|grasp|seize|snatch} your {rss|rss feed} as I {can not|can't} {in finding|find|to find} your {email|e-mail} subscription {link|hyperlink} or {newsletter|e-newsletter} service. Do {you have|you've} any? {Please|Kindly} {allow|permit|let} me {realize|recognize|understand|recognise|know} {so that|in order that} I {may just|may|could} subscribe. 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[reformata.net]

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Is this a paid theme or did you {customize|modify} it yourself? {Either way|Anyway} keep up the {nice|excellent} quality writing, {it's|it is} rare to see a {nice|great} blog like this one {these days|nowadays|today}.| {I am|I'm} {extremely|really} {inspired|impressed} {with your|together with your|along with your} writing {talents|skills|abilities} {and also|as {smartly|well|neatly} as} with the {layout|format|structure} {for your|on your|in your|to your} {blog|weblog}. {Is this|Is that this} a paid {subject|topic|subject matter|theme} or did you {customize|modify} it {yourself|your self}? {Either way|Anyway} {stay|keep} up the {nice|excellent} {quality|high quality} writing, {it's|it is} {rare|uncommon} {to peer|to see|to look} a {nice|great} {blog|weblog} like this one {these days|nowadays|today}..| {Hi|Hello}, Neat post. {There is|There's} {a problem|an issue} {with your|together with your|along with your} {site|web site|website} in {internet|web} explorer, {may|might|could|would} {check|test} this? IE {still|nonetheless} is the {marketplace|market} {leader|chief} and {a large|a good|a big|a huge} {part of|section of|component to|portion of|component of|element of} {other folks|folks|other people|people} will {leave out|omit|miss|pass over} your {great|wonderful|fantastic|magnificent|excellent} writing {due to|because of} this problem.| {I'm|I am} not sure where {you are|you're} getting your {info|information}, but {good|great} topic. I needs to spend some time learning {more|much more} or understanding more. Thanks for {great|wonderful|fantastic|magnificent|excellent} {information|info} I was looking for this {information|info} for my mission.| {Hi|Hello}, i think that i saw you visited my {blog|weblog|website|web site|site} {so|thus} i came to “return the favor”.{I am|I'm} {trying to|attempting to} find things to {improve|enhance} my {website|site|web site}!I suppose its ok to use {some of|a few of} your ideas!\

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{ {I have|I've} been {surfing|browsing} online more than {three|3|2|4} hours today, yet I never found any interesting article like yours. {It's|It is} pretty worth enough for me. {In my opinion|Personally|In my view}, if all {webmasters|site owners|website owners|web owners} and bloggers made good content as you did, the {internet|net|web} will be {much more|a lot more} useful than ever before.| I {couldn't|could not} {resist|refrain from} commenting. {Very well|Perfectly|Well|Exceptionally well} written!| {I will|I'll} {right away|immediately} {take hold of|grab|clutch|grasp|seize|snatch} your {rss|rss feed} as I {can not|can't} {in finding|find|to find} your {email|e-mail} subscription {link|hyperlink} or {newsletter|e-newsletter} service. Do {you have|you've} any? {Please|Kindly} {allow|permit|let} me {realize|recognize|understand|recognise|know} {so that|in order that} I {may just|may|could} subscribe. 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Keep on posting!| I {{leave|drop|{write|create}} a {comment|leave a response}|drop a {comment|leave a response}|{comment|leave a response}} {each time|when|whenever} I {appreciate|like|especially enjoy} a {post|article} on a {site|{blog|website}|site|website} or {I have|if I have} something to {add|contribute|valuable to contribute} {to the discussion|to the conversation}. {It is|Usually it is|Usually it's|It's} {a result of|triggered by|caused by} the {passion|fire|sincerness} {communicated|displayed} in the {post|article} I {read|looked at|browsed}. And {on|after} this {post|article} SLA Biomedical & Life Sciences Division Blog: Do Snake Venoms Evolve According to the Prey They Are Intended to Kill, Or Do Snakes Just Dial Up the Dosage or Concentration of Whatever Venom They Have to the Size of the Prey in their Sights?. I {{was|was actually} moved|{was|was actually} excited} enough to {drop|{leave|drop|{write|create}}|post} a {thought|{comment|{comment|leave a response}a response}} {: -P|:)|;)|;-)|:-)} I {do have|actually do have} {{some|a few} questions|a couple of questions|2 questions} for you {if you {don't|do not|usually do not|tend not to} mind|if it's {allright|okay}}. {Is it|Could it be} {just|only|simply} me or {do|does it {seem|appear|give the impression|look|look as if|look like} like} {some|a few} of {the|these} {comments|responses|remarks} {look|appear|come across} {like they are|as if they are|like} {coming from|written by|left by} brain dead {people|visitors|folks|individuals}? :-P And, if you are {posting|writing} {on|at} {other|additional} {sites|social sites|online sites|online social sites|places}, {I'd|I would} like to {follow|keep up with} {you|{anything|everything} {new|fresh} you have to post}. {Could|Would} you {list|make a list} {all|every one|the complete urls} of {your|all your} {social|communal|community|public|shared} {pages|sites} like your {twitter feed, Facebook page or linkedin profile|linkedin profile, Facebook page or twitter feed|Facebook page, twitter feed, or linkedin profile}?| {Hi there|Hello}, I enjoy reading {all of|through} your {article|post|article post}. I {like|wanted} to write a little comment to support you.| I {always|constantly|every time} spent my half an hour to read this {blog|weblog|webpage|website|web site}'s {articles|posts|articles or reviews|content} {everyday|daily|every day|all the time} along with a {cup|mug} of coffee.| I {always|for all time|all the time|constantly|every time} emailed this {blog|weblog|webpage|website|web site} post page to all my {friends|associates|contacts}, {because|since|as|for the reason that} if like to read it {then|after that|next|afterward} my {friends|links|contacts} will too.| My {coder|programmer|developer} is trying to {persuade|convince} me to move to .net from PHP. I have always disliked the idea because of the {expenses|costs}. But he's tryiong none the less. I've been using {Movable-type|WordPress} on {a number of|a variety of|numerous|several|various} websites for about a year and am {nervous|anxious|worried|concerned} about switching to another platform. 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